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matrix_synapse_storage_path is already defined in matrix-synapse/defaults/main.yml (with a default of "{{ matrix_synapse_base_path }}/storage"), but was not being used for its presumed purpose in matrix-synapse.service.j2. As a result, if matrix_synapse_storage_path was overridden (in a vars.yml), the synapse service failed to start.
recklesscoder